Bhṛgu–Bharadvāja-saṃvāda: Vānaprastha-parivrājaka-ācāra, Abhaya-dharma, and Lokānāṃ Vibhāga (Śānti-parva 185)
पुण्यापुण्यैस्तथा गन्धैर्धूपैश्व विविधैरपि । अरोगाः: पुष्पिता: सन्ति तस्माज्जिप्रन्ति पादपा:
puṇyāpuṇyais tathā gandhair dhūpaiś ca vividhair api | arogāḥ puṣpitāḥ santi tasmāj jighranti pādapāḥ ||
バラドヴァージャは言った。「清浄と不浄とされる香り、また種々の薫香の煙によって、樹々は病を離れ、花を開き実を結ぶ。これにより、樹にもまた嗅覚のはたらきがあると定まる。」
भरद्वाज उवाच
The verse argues by observable effect (health and blossoming due to fragrances and incense) that trees possess a sensory capacity—specifically smell—thus extending consideration of sentience beyond humans and animals.
In a didactic discussion in Śānti Parva, Bharadvāja presents an inference: since trees respond beneficially to scents and fumigation, it is reasonable to conclude that they can smell.