Bhṛgu–Bharadvāja-saṃvāda: Vānaprastha-parivrājaka-ācāra, Abhaya-dharma, and Lokānāṃ Vibhāga (Śānti-parva 185)
पुण्यापुण्यैस्तथा गन्धैर्धूपैश्व विविधैरपि । अरोगाः: पुष्पिता: सन्ति तस्माज्जिप्रन्ति पादपा:
puṇyāpuṇyais tathā gandhair dhūpaiś ca vividhair api | arogāḥ puṣpitāḥ santi tasmāj jighranti pādapāḥ ||
Bharadvāja dit : «Par des fragrances tenues pour pures et impures, et aussi par la fumée d’encens de maintes sortes, les arbres deviennent exempts de maladie et se couvrent de fleurs et de fruits. Par là, il est établi que les arbres aussi possèdent la faculté de l’odorat.»
भरद्वाज उवाच
The verse argues by observable effect (health and blossoming due to fragrances and incense) that trees possess a sensory capacity—specifically smell—thus extending consideration of sentience beyond humans and animals.
In a didactic discussion in Śānti Parva, Bharadvāja presents an inference: since trees respond beneficially to scents and fumigation, it is reasonable to conclude that they can smell.